Transcribed for String Orchestra the second movement of the New World Symphony is introduced by a harmonic progression of chords in the wind instruments. Beckerman interprets these chords as a musical rendition of the narrative formula "Once upon a time". Then a solo cor anglais (English horn) plays the famous main theme in D-flat major accompanied by muted strings. Dvořák was said to have changed the theme from clarinet to cor anglais as it reminded him of the voice of Harry Burleigh. The movement's middle section contains a passage in C♯ minor evoking a nostalgic and desolate mood which eventually leads into a funeral march above pizzicato steps in the basses. It is followed by a quasi-scherzo that incorporates this movement's theme as well as the first movement's main and closing themes. The Largo is concluded with the soft return of the main theme and introductory chords.

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